The Wilson Ornithological Society

Guide to Graduate Studies in Ornithology in North America

Compiled by the Committee on Undergraduate Outreach of the Wilson Ornithological Society
June 2003; updated May 2005

The purpose of this guide is to give undergraduate college students and their academic advisors information about those graduate degree programs where they can do advanced studies in support of their interests in the biology of birds. We present information abstracted from brochures and catalogs furnished by the departments and faculty members of the institutions listed below.

Please send corrections and additions to Douglas White, Chair <>

This guide organizes programs by geographic region. These regions are the following:

North East: Canada east of Manitoba, south to Ohio, Pennsylvania, and New Jersey.

South East: Kentucky, West Virginia, Maryland, and Delaware, south to Mississippi, Alabama, and Florida.

North Central: Saskatchewan and Manitoba south to Nebraska, Iowa, Illinois, and Indiana.

South Central: Kansas, and Missouri south to Texas, and Louisiana.

North West: Alaska, and North West Territories south to Oregon, Idaho, and Wyoming.

South West: California to Colorado, and south through Mexico.

Each entry in this guide takes the following form:

  • Name and address of the institution.
  • Description of the degree program.
  • Description of facilities supporting ornithological research.
  • Names and research interests of faculty who direct ornithological studies.
  • Addresses and telephone numbers to contact for further information.



The University of Connecticut, Storrs, CT 06269-3043.

Description.--The University of Connecticut, Storrs, located about 20 miles east of Hartford, has offered courses in ornithology since 1900, when the institution was called the Connecticut Agricultural College. Students may earn both M.S. and Ph.D. degrees in the Departments of Ecology and Evolutionary Biology, Physiology and Neurobiology, and Psychology; and the M.S. degree in the Department of Natural Resource Management and Engineering.

Facilities.--The University of Connecticut owns about 2300 acres, mostly undeveloped and available for field research. Besides this, many thousands of acres of nearby woodlands are readily available, and there is a marine field station at Noank, CT. The University of Connecticut Museum of Natural History contains more than 9000 study skins of birds, as well as skeletons, embalmed specimens, feathers, nests, eggs, and photographs.

Faculty.--Those faculty currently involved in ornithological studies are:

  • George A. Clark, Jr., Department of Ecology and Evolutionary Biology: Behavior, biogeography, evolution, and integumental structure of birds.
  • Andrew Moiseff, Department of Physiology and Neurobiology: hearing in Barn Owls.
  • David B. Miller, Department of Psychology: ontogeny of vocalization in ducks.
  • John Barclay, Department of Natural Resource Management and Engineering: Wildlife management and conservation of waterfowl.
  • Daniel L. Civco, Department of Natural Resource Management and Engineering: GIS and avian biogeography.
For information.--Write to Graduate Admissions, The University of Connecticut, Storrs, CT 06269-3043, or write to the department of interest.



The University of Maine, Orono, ME 04469.

Description.--The University of Maine emphasizes biological sciences and natural resources conservation, covered by eight departments (Wildlife Ecology; Plant Biology and Pathology: Forest Ecosystems; Forest Management; Applied Ecology and Environmental Sciences; Oceanography; Zoology; and Resource Economics and Policy) with about 30 faculty members who comprise a Conservation Biology Interest Group. The degree of Ph.D. may be earned in the fields of Biological Sciences, Ecology and Environmental Science, Plant Science, Wildlife Ecology, and Zoology. The Master of Science degree may be earned in Resource Economics; Botany and Plant pathology; Ecology and Environmental Science; Entomology; Plant, Soil, and Environmental Sciences; Resource Utilization; Wildlife Ecology; and Zoology.

Facilities.--The forest, wetland, freshwater, and marine ecosystems of Maine offer a diverse biota near campus for conservation biology research.

Faculty.--Those currently involved in ornithological studies are:

  • William E. Glanz, Department of Zoology: Community ecology of mammals and birds, foraging ecology and social behavior of granivorous mammals and birds, evolution and biogeography of North and South American rodents.
  • Malcolm L. Hunter, Jr., Department of Wildlife Ecology: Conservation biology, forest wildlife management, landscape ecology, international conservation, including effects of forest conditions on bird populations.
  • William B. Krohn, Department of Wildlife Ecology: Migratory bird management, habitat evaluation, wildlife administration.
  • Raymond J. O'Connor, Department of Wildlife Ecology: Bird population ecology, habitat dynamics, bioindicator dynamics.
  • Ray B. Owen, Jr., Department of Wildlife Ecology. Waterfowl and wetlands ecology, avian ecology, wildlife policy.
  • Frederick A. Servello, Department of Wildlife Ecology: Vertebrate nutrition and physiology, habitat relationships of birds and mammals.
For information.--Write to The Graduate School, Winslow Hall, University of Maine, Orono, ME 04469. To inquire about specific opportunities and the availability of graduate assistantships, write to any of the faculty members listed whose interests are close to yours. For general information about Conservation Biology, write to Malcolm Hunter, Department of Wildlife Ecology.



Antioch New England Graduate School, Keene, NH 03431-3516

Description. &endash; For more than thirty years, Antioch New England has provided leadership in the development of interdisciplinary graduate programs. The Masters Program, founded in 1972, was one of the first environmental studies programs in North America. Graduates and faculty form an influential network of professionals throughout the Northeast. The program stresses small classes, high quality advising and mentoring, and non-traditional, participatory learning approaches. The Doctoral program, founded in 1995, is designed for the experienced environmental professional and offers an interdisciplinary, research-oriented Ph.D.

Like many environmental study programs, both Masters and Doctoral programs emphasize a broad body of knowledge including science, policy, natural history, and ethics. But unlike most graduate programs, we are distinguished by two important components&emdash;our applied focus and reflective practice. Students develop critical academic skills through practical application and applied scholarship. They develop professional and scholarly networks that are directly engaged with schools, community groups, and environmental organizations or agencies. We also believe that effective environmental scholars and professionals must be engaged in reflective practice. It is vital that we are capable of articulating and projecting our values&emdash;what we describe as ecological identity. The M.S. degree may be earned in the Department of Environmental Studies with specialization in Conservation Biology, Resource Management and Administration, Environmental Education, and Teacher Certification in Biology or General Science.

Faculty.--The faculty members currently supporting research specifically with birds are:

  • Jonathan L. Atwood: Population dynamics of neotropical migrant songbirds; role of rare and threatened bird species as focal elements of regional conservation planning; factors influencing site selection and fidelity of terns.

    For information.&endash; Write the Environmental Studies Department, Antioch New England Graduate School, 40 Avon Street, Keene, NH 03431-3516; telephone (603) 357-3122, FAX: (603) 357-0718. Information also available on the WWW:


Dartmouth College, Hanover, NH 03755-3576

Description.--Dartmouth College has an undergraduate student body numbering about 4300, and a graduate student body of about 850. Hanover is a town in the heart of the New England winter skiing and summer lake resort areas, about a two-hour drive from Boston. The degree of Ph.D. may be earned in the Department of Biology in the fields of Ecology and Evolutionary Biology, and Molecular Biology.

Facilities.--The Charles Gilman Life Sciences Laboratory provides 62500 square feet of research and teaching facilities. This includes three cold rooms, one sub-zero room, eight constant environment rooms, a greenhouse, and quarters for maintenance of cold-blooded animals. Animal care facilities for warm-blooded animals are available in the adjoining medical school research building. There are tissue culture rooms, extensive light microscopy facilities, and a facility for both transmission and scanning electron microscopy. Various other standard kinds of research equipment are available. Biological field investigators have ready access to a variety of habitats and ecosystems, including many small, clear streams, lakes, ponds, bogs, the Connecticut River (on campus), boreal forests, deciduous forests, and alpine tundra. Established research sites are maintained at numerous locations in New England, including a 26000 acre tract in northern New Hampshire with two rivers, and the Hubbard Brook Experimental Forest in the White Mountains. There are also research sites in Jamaica, Costa Rica, and Indonesia.

Faculty.--The faculty members currently supporting research specifically with birds are:

  • Richard T. Holmes: Mating systems, reproductive ecology, territoriality, site fidelity and dispersal, survivorship, habitat selection, foraging behavior, and related population attributes of birds. Emphasis is on migratory songbirds in both their temperate zone breeding areas and their tropical winter quarters.
  • Douglas T. Bolger: Conservation biology, population ecology, and habitat requirements of birds, with emphasis on the effects of habitat fragmentation.
For information.--Write the Chair, Graduate Admissions Committee, Department of Biological Sciences, 6044 Gilman, Dartmouth College, Hanover, New Hampshire 03755-3576, telephone (603) 646-3847, FAX: (603) 646-1347. Information also available on the WWW:



Cornell University, Ithaca, NY 14853.

Description.--Although Cornell University does not offer a degree specifically in ornithology, it has long been known as a major center of ornithological research. Dr. Arthur A. Allen was appointed one of the nation's first professors of ornithology in 1917. Since then, Cornell has become a famous center for study of avian biology. The degree of Ph.D. may be earned in several fields, and departments. Those that provide opportunities for study with birds include Anatomy; Ecology and Systematics; Natural Resources; Neurobiology and Behavior; Psychology; and Veterinary Medicine.

Facilities.--The Cornell Laboratory of Ornithology is a world-famous center for the study, appreciation, and conservation of birds. It houses the Library of Natural Sounds, holding one of the largest collections of bird sound recordings in the world. The Bioacoustics Research Program provides advanced facilities for bird sound analysis. General biological research facilities are extensive and varied. Laboratories house equipment and instrumentation for molecular, physiological, behavioral, and environmental studies. Four thousand acres of Cornell farmland and forest as well as 30000 acres of New York State lands are within easy reach of the campus. Field laboratories are maintained on Cayuga and Oneida Lakes, and in the Adirondack Mountains. The Section of Ecology and Systematics maintains a bird collection representing about half the known species of birds of the world, and nearly all avian families.

Faculty.--Those faculty members supporting ornithological studies are:

  • Elizabeth Adkins-Regan, Section of Neurobiology and Behavior and Department of Psychology: Reproductive behavior of birds, especially endocrine and neural mechanisms of sexual behavior.
  • Christopher W. Clark, Laboratory of Ornithology: Director of the Bioacoustics Research Program, investigating animal acoustic communication and developing acoustic technologies with applications in conservation and environmental monitoring. Studies acoustic networks within territorial songbird communities, evolutionary role and selective advantages of countersinging, and develops techniques to augment avian census efforts.
  • Timothy DeVoogt, Department of Psychology: Neural basis for song acquisition and expression in songbirds, including hormonal influences and brain changes responsible for song learning and other behaviors.
  • AndrJ Dhondt, Laboratory of Ornithology: Director of Bird Population Studies program, he combines various disciplines such as population dynamics, behavioral ecology, and quantitative genetics in long-term field studies of individually marked birds.
  • Stephen T. Emlen, Section of Neurobiology and Behavior: Studies the adaptive significance of the diverse social organizations of birds and mammals, including evolution of cooperative breeding and helping at the nest, polygynous and polyandrous mating systems, male-female conflicts of interest within various mating systems, and patterns of parental investment. He also maintains research interests in animal orientation and navigation, and avian acoustical communication.
  • Howard E. Evans, College of Veterinary Medicine: avian anatomy, and anatomical illustration.
  • Thomas A. Gavin, Department of Natural Resources: Ecology, behavior, and conservation of birds, using genetic techniques to investigate the relationships among breeding populations.
  • Howard C. Howland, Section of Neurobiology and Behavior: Physiology of vision, to elucidate mechanisms of focusing and regulation of growth in avian eyes.
  • George V. Kollias, College of Veterinary Medicine: Avian pathobiology, immunology/cell biology, and pharmacology.
  • Richard A. Malecki, Department of Natural Resources: Waterfowl biology and wetland ecosystems.
  • Kevin J. McGowan, Section of Ecology and Systematics. Social and reproductive behavior in crows and jays, and he is the principal caretaker of the Cornell ornithology collection.
  • Milo E. Richmond, Department of Natural Resources: Vertebrate ecology and reproduction, with studies of a wide range of avian types.
  • Richard B. Root, Section of Ecology and Systematics: Foraging ecology, biogeography, and community relationships of birds.
  • Paul W. Sherman, Section of Neurobiology and Behavior: Behavioral ecology of many avian species, the causes and consequences of extra-pair copulations in passerines, and the costs and benefits of extreme brood parasitism (dump nesting) in hole-nesting ducks.
  • Charles R. Smith, Department of Natural Resources: Conservation of North American bird species at the population, community, and landscape levels.
  • Charles Walcott, Laboratory of Ornithology: Director of the Laboratory of Ornithology, his main interests are bird navigation, orientation, and acoustic communication.
  • David W. Winkler, Section of Ecology and Systematics: Evolutionary and behavioral ecology of birds, focusing on the phylogeny and physiological and ecological determinants of life history traits.
For information:--Contact the Department, Section, or faculty member whose activities match your interests. The Addresses of the departments and sections are:

Cornell Laboratory of Ornithology: Education and Information Services, Cornell Laboratory of Ornithology, 159 Sapsucker Woods Road, Ithaca, NY 14850, (607) 254-2440.

Section of Ecology and Systematics: Cornell University, Section of Ecology and Systematics, E145 Corson Hall, Ithaca, NY 14853-2701, (607) 255-4522.

Section of Neurobiology and Behavior: Cornell University, Section of Neurobiology and Behavior, W363 Seeley G. Mudd Hall, Ithaca, NY 14853-2702, (607) 255-4517.

Department of Natural Resources: Cornell University, Department of Natural Resources, 118 Fernow Hall, Ithaca, NY 14853-3001, (607) 255-2821.

Department of Psychology: Cornell University, Department of Psychology, 211 Uris Hall, Ithaca, NY 14853-7601, (607) 255-3834.

College of Veterinary Medicine: Cornell University, New York State College of Veterinary Medicine, Ithaca, NY 14853-6401, (607) 253-3000.


University at Albany, State University of New York, Albany, NY 12214-0222.

Description.--The campus is located two miles from downtown Albany, a city of 100,000 near Saratoga, Schenectady, and Troy. The degrees of M.S. and Ph.D. can be earned in the Department of Biological Sciences in the specializations of cellular and developmental biology, ecology and animal behavior, molecular biology, and neurobiology. Each specialization has separate examinations and curricular requirements.

Facilities.--The Biological Sciences Building has nearly three acres of space containing numerous research laboratories equipped for modern research. Among the facilities are scanning and transmission electron microscopes, aquarium room, small animal facility, and numerous environmental chambers. Students in ecology and animal behavior have nearby access to off campus research facilities at the Albany Pinebush and Huyck Preserves.

Faculty.--The faculty members supporting studies in ornithology are:

  • Kenneth P. Able, Ecology and Animal Behavior: Animal migration, orientation, and navigation.
  • Jerram L. Brown, Ecology and Animal Behavior: Social behavior; behavioral ecology; evolutionary biology; major histocompatibility complex; mate choice.
  • Caro-Beth Stewart, Molecular Biology: Molecular basis for adaptive evolution; evolution of digestive enzymes; evolution of substrate specificity; tree analysis of gene families.
For information.--Contact Graduate programs Secretary, Department of Biological Sciences, University at Albany, State University of New York, 1400 Washington Avenue, Albany, NY 12222, Tel. (518) 442-4300, FAX (518) 442-4354. Visit the Biological Sciences Department web site at



The Ohio State University, Columbus, OH 43210

Description.--The Department of Zoology offers the degree of Ph.D., and enrolls more than 60 graduate students, many of whom are engaged in studies of avian biology. Faculty members are involved in such areas of avian biology as behavior, conservation, ecology, evolution, genetics, physiology, social systems, and vocal communication.

Facilities.--The Department of Zoology provides laboratories for genetics, electrophysiology, and general research. Libraries for Biology and Pharmacy, Agriculture, Veterinary Medicine, as well as the Main Library offer access to many research resources. The Museum of Biodiversity houses the Museum of Zoology, containing avian skeletons, study skins, and eggs, plus the Trautman/Condit Collection, a small private library with many rare publications. Also in the Museum is the Borror Laboratory of Bioacoustics, one of the major facilities for the study of avian vocalizations. The Ohio Cooperative Fish and Wildlife Research Unit, funded by the U.S. Department of the Interior, is located in the Department of Zoology. Its funding underwrites many student projects. The Department of Zoology also is closely associated with the Department of Natural Resources, which has several ongoing projects involving endangered and threatened bird species. The Veterinary School also offers opportunities with exotic species and avian surgical techniques.

Faculty.--Those involved in research in avian biology are:

  • Richard A. Bradley: Biology of vocal behavior in birds, emphasizing the evolutionary significance of geographic variation in bird song, female song in passerines.
  • Sandra L. L. Gaunt: Biology of vocal behavior in animals, with emphasis on behavioral ecology, evolution of vocal behavior, and vocal learning in avian taxa.
  • Thomas C. Grubb, Jr.: Theoretical and empirical behavioral ecology and conservation biology.
  • Douglas A. Nelson: Communication behavior and song development in birds.
  • Patricia G. Parker: Ecology and evolution of social behavior and the application of principles of population biology to wildlife conservation.
  • Joseph B. Williams: Avian physiology with emphasis on field metabolic rates and water flux of free-living birds.
For information.--Department of Zoology, The Ohio State University, 1735 Neil Avenue, Columbus, OH 43210-1293. Telephone (614) 292-8088, FAX (614) 292-2030.



The Pennsylvania State University, University Park, PA 16802-4703

Description.--Penn State University offers M.S. and Ph.D. degrees in both Wildlife and Fisheries Science, and Ecology. In both programs, students are involved in graduate research with birds. The Wildlife and Fisheries Science Program is in the School of Forest Resources. Graduate students in this program study a variety of topics including avian responses to altered ecosystems, human impacts on bird populations, and the ecology and habitat requirements of non-game birds. The Ecology Program is an intercollege graduate degree program designed to teach students both basic and applied aspects of ecology. Students conduct research on birds similar to that described for the Wildlife and Fisheries Science Program. Course work differs between the two programs, with a greater emphasis on basic ecology in the Ecology Program.

Facilities.--A small museum collection and collaborative research with Hawk Mountain Sanctuary.

Faculty.--Those currently active in studies on biology of birds are:

  • Margaret C. Brittingham, Associate Professor of Wildlife Resources: avian ecology, human impacts on bird populations, breeding ecology of neotropical migrants, urban birds.
  • Robert P. Brooks, Associate Professor of Wildlife Ecology: wetland ecology, avian use of wetlands.
  • Richard H. Yahner, Professor of Wildlife Conservation: forest wildlife, neotropical migrants.
For information.--About the Wildlife and Fisheries Science Program, write to: Assistant Director for Research and Graduate Studies, School of Forest Resources, 213 Ferguson Building, The Pennsylvania State University, University Park, PA 16802.

About the Ecology Program, write to: Richard H. Yahner, Chair, IGDP in Ecology, 107 Ferguson Building, Pennsylvania State University, University Park, PA 16802.


Villanova University, Villanova, PA 19085-1699

Description.--Villanova University is a Roman Catholic university sponsored by the religious order of St. Augustine. It is situated in Villanova, Pennsylvania, 12 miles west of Philadelphia. The Department of Biology offers a program of study leading to the Master of Science degree in biology (with thesis) or the Master of Arts degree in biology (without thesis) ), and a combined 5-year B.S./M.S. degree in Biology (with thesis). Ornithology is one of the areas in which thesis research may be undertaken.

Facilities.--Laboratories are housed in a modern, air-conditioned building (completely renovated and expanded in 2000), equipped for graduate instruction and research in many areas of biology. There is a greenhouse, an animal-care facility, darkroom, electron microscopes (TEM, SEM), instruments for work with radioisotopes, X-ray facility, cell culture laboratory, three automated DNA sequencers, a Geographic Information System (GIS) running ArcView and ARC/INFO software, and various research instruments and computation facilities. Opportunities for ornithological field study include collaborative programs involving Hawk Mountain Sanctuary Association (Kempton, PA) and Archbold Biological Station (Lake Placid, FL).

Faculty.--Those currently involved in ornithological research are:

  • Robert L. Curry: Behavioral ecology, focusing on vertebrate social organization; ornithology, conservation biology. Social ecology of Florida Scrub-Jays; hybridization in Black-capped and Carolina Chickadees; conservation ecology of island-endemic mockingbirds and allies.
  • John M. Olson: Metabolic and muscle physiology; ecological physiology, development and physiology of endothermy in birds.
For Information.--Write Chairman, Department of Biology, Villanova University, Villanova, Pennsylvania 19085. Tel. (610) 519-4830, FAX (610) 519-7863. E-mail:;



McGill University, Macdonald Campus, 21,111 Lakeshore Road, Ste-Anne-de-Bellevue, Quebec, Canada H9X 3V9

Description.--Graduate studies in Wildlife Biology have been conducted in the Department of Natural Resource Sciences at the Macdonald Campus since the late 1960s. From the beginning there has been an emphasis on ecological aspects conducted from often remote field locations. However, with the recent establishment of the Avian Science and Conservation Centre on the campus, studies also now involve captive animals. The emphasis is on practical application and is aimed at conservation of natural resources.

Facilities.--The campus holds the Avian Science and Conservation Centre, and the Ecomuseum of the St. Lawrence Valley Natural History Society, where excellent captive bird holding facilities are available.

Faculty.--Those supervising studies of birds are:

  • David M. Bird: Director, Avian Science and Conservation Centre, research on birds in a variety of areas including reproductive physiology, toxicology, genetics, parasitology, behavioral ecology, nutrition, physiology, and wildlife management.
  • Rodger D. Titman: Behavior and ecology of waterfowl and wetland ecology; social systems, behavioral ecology, and habitat selection by ducks and other birds.
  • K. A. Blanchard: Executive Vice-President, Quebec-Labrador Foundation is an adjunct professor involved in environmental education and conservation of seabirds.
  • Henry Markon: Wetland ecologist with Institute for Waterfowl & Wetland Research, Ducks Unlimited, is an adjunct professor studying waterbirds in wetlands.
  • Jean-Pierre Savard: Research scientist with the Canadian Wildlife Service, is an adjunct professor who studies waterfowl and forest bird ecology, behavior, and management.
For information.--Contact Graduate Program Secretary, Department of Natural Resource Sciences, McGill University, Macdonald Campus, 21,111 Lakeshore Road, Ste. Anne de Bellevue, Quebec, H9X 3V9, Canada. Tel. (514) 398-7941, FAX (514) 398-7990




Auburn University, Auburn, AL 36849

Description.--The Department of Zoology and Wildlife Science at Auburn University trains students in its Program in Field Ornithology. The department offers M.S. and Ph.D. degrees with specialties in Zoology and in Wildlife Science. Students in the Department investigate a wide range of topics in fields as diverse as behavioral ecology, physiological ecology, evolutionary ecology, community ecology, population ecology, landscape ecology, and wildlife ecology. Areas of faculty specialization include sexual selection and the evolution of avian plumage coloration; ecology of tropical bird communities; management of waterfowl, raptor, and game species; and reproductive ecology of waterfowl and songbirds. Additional topics are pursued by faculty and graduate students in the Department. Visit the Department's web page (see below) for more information.

Facilities.--Resources specifically supporting research on birds are a dove research laboratory with wet lab, computer facilities, library, incubation equipment, and outdoor holding and breeding facilities; an avian research center with 25 large outdoor aviaries, live bird room, wet lab, enclosures for mate choice experiments, and computers; the Alabama Cooperative Fish and Wildlife Research Unit, which provides administrative support for student research, computer laboratory, and field equipment and vehicles; and the Southeastern Raptor Rehabilitation Center.

Faculty.--Faculty members and research associates currently studying birds include:

  • James Armstrong, Professor: Wildlife damage management, especially bird predators at aquaculture ponds. (e-mail:
  • Barry Grand, Associate Professor; Leader, Alabama Cooperative Fisheries and Wildlife Research Unit: Productivity and management of acrtic and subarctic breeding waterfowl; population dynamics; factors influencing productivity and survival; study of marked populations. (e-mail:
  • Gary Hepp, Professor: Ecology and management of waterfowl populations; reproductive ecology of Wood Duck; winter ecology of waterfowl; effects of herbivory by waterfowl and other large herbivores on aquatic plant communities. (e-mail:
  • Geoffrey Hill, Associate Professor: Sexual selection and mate choice in cardueline finches; avian mating systems; carotenoid plumage pigmentation; habitat use by passerine birds; management of Neotropical migrant birds. (e-mail:
  • Ralph Mirarchi, Professor: Raptor rehabilitation and management; dove and pigeon ecology. (e-mail:
  • Tara R. Robinson, Research Associate: Behavioral ecology, emphasizing evolution of families, social behavior, and cooperative breeding. (e-mail:
  • W. Douglas Robinson, Assistant Professor: Ecology of tropical bird communities; community ecology; neotropical migrant birds; landscape ecology and biogeography. (e-mail:
  • Lee Stribling, Professor: Bobwhite Quail management. (e-mail:
For information.--Department of Zoology and Wildlife Science, 331 Funchess Hall, Auburn University, Auburn, AL 36849. Tel. (334) 844-4850, FAX (334) 844-9234.



The University of Georgia, Athens, GA 30602.

Description.--The University of Georgia, chartered in 1785 enrolls approximately 25000 students, about 5000 of whom are graduate students. Athens is located in the Piedmont Region of northeastern Georgia, a few hours drive from the Appalachian and Blue Ridge Mountains, and from the Atlantic and Gulf Coastal Plains. The School of Forest Resources offers graduate studies in ornithology and a wide variety of other wildlife-related subjects, leading to the degrees of M.S. and Ph.D. The M. S. degree is designed for students who plan to specialize in a particular area of wildlife or plan to enter a Ph.D. program, and includes a thesis as part of the requirements. The Ph.D. degree program is for students with an interest in original research in specific areas of wildlife.

Facilities.--The wide range of facilities for wildlife research available in the School of Forest Resources and other administrative units include general laboratories, computational services, and research libraries. Other support units include Biological Sciences, Pharmacy, Veterinary Medicine, the Institute of Ecology, the Institute of Natural Resources, the Southeastern Cooperative Wildlife Disease Study of the College of Veterinary Medicine, the Southeast Field Station of the U.S. Fish and Wildlife Service, and the Georgia Department of Natural Resources. There is also a close working relationship with the Savannah River Ecology Laboratory near Augusta. The Cooperative Fish and Wildlife Research Unit of the U. S. Fish and Wildlife Service is an integral part of the wildlife program.

Faculty.--Faculty members available to direct ornithological studies are the following:

  • John Avise, Genetics Department: Molecular ecology and evolution.
  • Lehr Brisbin, Senior Research Scientist, Savannah River Ecology Laboratory: Vertebrate ecology, ornithology, radioecology, behavioral ecology, studies of mathematical models of growth and contaminant cycling processes, studies of the ecology of domestication, ecotoxicology.
  • Brian R. Chapman, School of Forest Resources: Non-game ecology, endangered species management, vertebrate natural history.
  • Michael J. Conroy, School of Forest Resources, Georgia Cooperative Fish and Wildlife Research Unit: Population dynamics, avian and mammalian population ecology, wildlife biometrics.
  • Patricia Adair Gowaty, Institute of Ecology: Behavioral and evolutionary ecology, mating systems, sex allocation, statistical aspects of genetical parentage testing.
  • David G. Krementz, U. S. Fish and Wildlife Service, Southeast Research Group, and School of Forest Resources: Population ecology, upland game bird biology, waterfowl ecology.
  • Ernest E. Provost, School of Forest Resources; Wildlife biology, vertebrate natural history.
  • Ron Pulliam, Institute of Ecology: Ecology and behavior of birds; species diversity and population ecology.
  • Sara H. Schweitzer, School of forest Resources; Effects of human disturbance on ecosystems and wildlife populations, conservation biology, ecological systems analysis, relationships among wildlife habitat selection, nutrition, and productivity; conservation of threatened and endangered species and ecosystems.
  • Donald H. White, School of Forest Resources, and Southeast Field Station, Patuxent Wildlife Research Center, U. S. Fish and Wildlife Service: Biocontaminants and avian ecology.
For information.--Write to Graduate Program, Wildlife Ecology and Management, School of Forest Resources, University of Georgia, Athens, GA 30602.


Georgia Southern University, Statesboro, GA 30460.

Description.-- Georgia Southern University is a comprehensive public university that enrolls approximately 16,000 students. Located on the coastal plain near Savannah, the campus provides easy access to some of the most biodiverse habitats in the southeast. The Department of Biology offers a strong MS program that provides the opportunity for graduate study in ornithology and many other fields of biology as well. A strong faculty of 28 scientists directs a program whose graduates have been highly successful. Recent ornithology graduates now study in top PhD programs, work in state and federal agencies, teach, and are employed in environmental consulting.

Facilities.--The Department of Biology is housed in 2 buildings that offer laboratories, classrooms, animal rooms, and computer facilities. The department is also closely affiliated with the Institute for Arthropodology and Parasitology and the Applied Coastal Research Laboratory located at the Skidaway Institute for Oceanography. Faculty and graduate students also have close working relationships with the St. Catherines Island Species Survival Center, Fort Stewart, the Georgia Department of Natural Resources, Gray's Reef Marine Sanctuary, Sapelo Island National Estuarine Research Reserve, Mountain Lake Biological Station, and the Caribbean Marine Research Center.

Faculty.--Faculty members available to direct ornithological studies are the following:

  • C. Ray Chandler: Ecology, behavior, and conservation of birds and other terrestrial vertebrates. (
  • Lance A. Durden: Systematics, ecology, and vector competence of ectoparasites of vertebrates, including birds. (
  • John W. Parrish: Comparative physiological ecology; life history of Southeastern American Kestrels. (

    For information.--Contact the Graduate Program Director, Department of Biology, Georgia Southern University, Statesboro, GA 30460-8042. Phone: 912-681-5487, Fax: 912-681-0845.



Towson University, Towson, MD 21252.

Description.--Towson University is part of the University of Maryland System, and is located in a northern suburb of Baltimore. The Department of Biology has 26 full-time faculty members, and approximately 60 graduate students. Students may earn the degree of M.S.

Facilities.--Abundant research space and an indoor aviary are available to graduate students. Research sites include several large wooded preserves in the surrounding area. Faculty conduct research at field stations in the mountains of western Maryland, and in north-central Wyoming.

Faculty.--Those active in ornithological studies are:

  • Don C. Forester: Behavioral ecology of vertebrates, especially parental care, sexual selection, and communication.
  • L. Scott Johnson: Mating systems and other reproductive behavior in birds, function of bird song, effects of ectoparasites on breeding success, ecology and conservation of secondary cavity nesting birds.
  • Brian S. Masters: Molecular aspects of ecology, evolution and behavior, especially parental care, kin recognition, sex ratios.
  • Erik P. Scully: Behavioral ecology, especially mechanisms of intraspecific competition for resources.
For information.--General information about the graduate program is available from Director of Graduate Admissions, The Graduate School, Towson University, Towson, MD 21252, telephone (410) 830-2500. For information of conducting ornithological research, contact Dr. L. Scott Johnson, Department of Biology, Towson University, Towson, MD 21252, telephone (410) 830-2587, E-mail:



Appalachian State University, Boone, NC 28608.

Description.--Appalachian State University is in the Blue Ridge of the Appalachian Mountains close to the borders of Virginia and Tennessee, less than two hours away from the region’s major airports and population centers. The University occupies a 75-acre main campus. The Biology Department offers the graduate degrees of M. S. in Biology, M. A. in Biology with Teacher Preparation. There are about 30 students currently enrolled in Master’s degree programs in Biology. The course offerings emphasize ecological and organismal approaches. Thesis research may be done in various aspects of ornithology.

Facilities.--Located in the Appalachians, Appalachian State offers ready access to study areas along the Blue Ridge Parkway, Grandfather Mountain (an International Biosphere Reserve), and Pisgah and Nantahala National Forests. Mount Mitchell State Park, the highest point east of the Mississippi, is 45 minutes away, and Great Smoky Mountains National Park is 3 hrs away. Faculty maintain working relationships with the National Park Service, US Fish and Wildlife Service, US Forest Service, and North Carolina’s nongame program. The University has its own 100-acre Environmental Studies Area, as well. There is a small teaching collection of bird skins, and an herbarium and extensive collection of amphibians and reptiles.

Faculty.--There are 22 faculty in the Biology Department, but only one of them is engaged in ornithological research with graduate students:

  • Matthew P. Rowe: Behavioral ecology and conservation biology, currently studying boreal relict birds species in the high-elevation spruce-fir forests of the southern Appalachians.
For information.--For general information about the graduate program in Biology, contact the Department Chair, telephone (828) 262-3025, Department of Biology, P. O. Box 32027, Boone, NC 28608-2027. Those interested specifically in ornithological studies may telephone Dr. Rowe at (828) 262-2676, or e-mail:


The College of William and Mary, P.O. Box 8795, Williamsburg, Virginia 23187-8795

Description.--The College of William & Mary is one of the two premier public universities in Virginia. It is also the second oldest university in North America and is regularly ranked as the best small public university in the United States. The Department of Biology offers a 2-year M.A. for those studying any aspect of ornithology. A joint Ph.D. is possible for students doing applied research (through the Program in Applied Science) or those doing marine ornithology (through the associated Virginia Institute of Marine Science).

Facilities.--William and Mary's location, near the shores of the Chesapeake Bay, opens up countless possibilities for field research on breeding colonial waterbirds or landbirds and raptors concentrated by this important migration bottleneck. The Biology Department's field house on the Eastern Shore National Wildlife Refuge allows convenient access to the lower bay. A field station run by the Virginia Institute of Marine Science allows access to the upper bay. The large and expanding aviary complex on campus is the cornerstone of the Biology Department's avian biology facilities. The immediate proximity to Colonial Historical National Park and large areas of publicly-owned wildlands creates numerous convenient study sites, including many acres of second-growth woodland owned by the Biology Department right on campus. All standard modern molecular and cellular biology laboratory equipment and scanning and transmission electron microscopes are available in common departmental facilities. The Center for Conservation Biology on campus has carried out hundreds of studies on endangered and threatened birds and is rapidly becoming one of the foremost bird conservation institutes in the mid-Atlantic region.

Faculty.--Those currently doing research with birds in the Biology Department are:

  • Ruth A. Beck, Conservation of colonial nesting waterbirds
  • Daniel A. Cristol, Migration, optimal foraging, spatial memory, wetland conservation
  • Mark H. Forsyth, Feather-degrading bacteria
  • John P. Swaddle, Communication, sexual selection, fluctuating asymmetry
  • Bryan D.Watts, Conservation, landscape ecology, foraging ecology
  • (Emeritus) Mitchell A. Byrd, Peregrine falcon reintroduction, eagle conservation

    For information contact Daniel A. Cristol, Department of Biology, P.O. Box 8795, College of William & Mary, Williamsburg, VA 23187-8795, phone: (757) 221-2405; FAX: (757) 221-6483; email:



Southern Illinois University, Carbondale, IL 62901-6501.

Description.--Ornithological studies in the Department of Zoology lead to Master of Science and Doctor of Philosophy degrees. A wide range of topics is supported by faculty expertise, including habitat use, foraging behavior, behavioral energetics, endangered species, predatory behavior, molecular evolution, toxicology, population dynamics, and various aspects of communication. Numerous parks and wildlife refuges are accessible in the southern Illinois region, which lies within the Mississippi Flyway.

Facilities.--In addition to the varied habitats and wildlife refuges available for field research sites in the vicinity of the university campus, SIUC provides laboratory space, computer facilites, electron microscopy facilities, vivarium, extensive main library, a collection of bird specimens, and a variety of field and laboratory equipment.

Faculty.--Those who supervise students working on avian research are:

  • Lee C. Drickamer: Animal behavior, behavioral ecology.
  • Robert J. Gates: Wildlife ecology, waterfowl and wetland management.
  • Richard S. Halbrook: Wildlife toxicology, population dynamics, and ecology.
  • Carey Krajewski: Molecular evolution, phylogenetics.
  • Jonathan Newman: Population dynamics and community ecology.
  • George H. Waring: Animal behavior, applied ethology, vertebrate natural history.
For information.--Contact any of these individuals by writing to them at the Department of Zoology, Southern Illinois University, Carbondale, IL 62901-6501. For forms for admission and financial aid, contact The Director of Graduate Studies, at the Department of Zoology, Telephone (618) 536-2314.


Illinois State University, Normal, IL 61790-4120.

Description.--The Department of Biological Sciences offers opportunities to earn the M.S. (thesis), and Ph.D. (dissertation) degrees based on research in avian behavior, biogeography, conservation biology, ecology, and systematics in both neotropical and nearctic birds. Students working on birds are encouraged to take advantage of other opportunities, including the M.S. (thesis) in Conservation Biology Sequence, and faculty in related areas of behavior, conservation biology, ecology, and evolution.

Facilities.--Facilities include molecular biology laboratories, research collection of skins and frozen tissues, and nearby field sites that include over 900 nestboxes for secondary cavity-nesting birds and hectares of on-going ecological restoration.

Faculty.--Those who supervise students working on avian research are:

  • Dr. Angelo Caparella: systematics, biogeography, conservation biology.
  • Dr. Charles F. Thompson: population ecology, behavior.
For information.--Contact any of these individuals directly or write Graduate Programs, Department of Biological Sciences, Illinois State University, Normal, IL 61790-4120. Telephone (309) 438-3669



Indiana University, Bloomington, IN 47405.

Description.--The Departments of Biology and Psychology and the Programs in Neural Science and Medicinal Sciences provide broad and intensive training for the Ph.D. degree in the in aspects of avian biology through the NSF-supported Center for the Integrative Study of Animal Behavior.

Facilities.--The wide range of faculty interests and departmental representation provide extensive facilities for studies in the topics of mate choice, sexual behavior, ecology of learning, neural plasticity, communication, orientation and migration, parental behavior, and development.

Faculty.--For information on faculty, write for information from the Department of Biology or The Department of Psychology.

For information.-- Contact Dr. Ellen D. Ketterson, Department of Biology, Jordan Hall, Indiana University, Bloomington, IN 47405, telephone (812) 855-6837, or Dr. William D. Timberlake, Department of Psychology, Psychology Building, Indiana University, Bloomington, IN 47405, telephone (812) 855-6837.



Iowa State University, Ames, Iowa 50011.

Description.&emdash; Iowa State University is a major research university located about 35 miles north of Des Moines. The M.S. and Ph.D. degrees are offered in the Department of Animal Ecology, the Department of Zoology and Genetics, and the interdepartmental Ecology and Evolutionary Biology program. The Department of Animal Ecology is also the home of two units with federal employees, the Iowa Cooperative Fish and Wildlife Research Unit and the Wildlife Habitat Management Institute.

Facilities.&emdash; The Department of Animal Ecology ( has a modern computer laboratory for graduate students, a Geographic Information Systems facility, a small ornithological teaching collection, laboratory facilities, and 14 aviaries. The Department of Zoology and Genetics ( has laboratory facilities for physiological and genetic studies.

Faculty.&emdash; Faculty members with expertise in some aspect of avian biology include:

  • Louis B. Best, Dept. Animal Ecology: Avian ecology, with particular emphasis on agroecosystems (
  • Bonnie S. Bowen, Dept. Animal Ecology: Ecology, evolution, and genetics (
  • William R. Clark, Dept. Animal Ecology: Population ecology and modeling of vertebrates, including waterfowl and Ring-necked Pheasant (
  • James J. Dinsmore, Dept. Animal Ecology: Community composition, habitat requirements, and colonization rates of birds on restored and natural wetlands (
  • William L. Hohman, Dept. Animal Ecology and Wildlife Habitat Management Institute (NRCS): Life history strategies of birds, especially waterfowl, and wildlife responses to land management practices (
  • Rolf R. Koford, Dept. Animal Ecology and Iowa Cooperative Fish and Wildlife Research Unit (USGS): Ecology, behavior, and population biology of birds that nest in grasslands, including songbirds and dabbling ducks (
  • David L. Otis, Dept. Animal Ecology and Iowa Cooperative Fish and Wildlife Research Unit (USGS): Demographic modeling of mourning doves and other species (
  • Tamara Stafford, Dept. Animal Ecology: Avian ecotoxicology (
  • Carol M. Vleck, Department of Zoology and Genetics: Physiological ecology, drawing on the disciplines of physiology, endocrinology, behavior, ecology and evolution (

    For information.&emdash; Contact one of the following offices or the faculty member whose activities match your interests.

    Department of Animal Ecology, Graduate Studies, 124 Science II, Iowa State University, Ames, IA 50011-3221, (515) 294-6148, FAX (515) 294-7874,

    Department of Zoology & Genetics, 1210 Molecular Biology Building , Iowa State University, Ames, IA 50011-3260, (515) 294-3909, FAX: (515) 294-6755,

    Ecology & Evolutionary Biology Graduate Program, 2018 Molecular Biology Building, Iowa State University, Ames, IA 50011, (515) 294-2196, FAX (515) 294-6790,



The University of Manitoba, Winnipeg, Manitoba, Canada R3T 2N2.

Description.--The University of Manitoba is the largest degree granting institution in Manitoba, and the oldest one in western Canada. Studies in ornithology may lead to the M. A. and Ph.D. degrees in Psychology and Zoology. The Psychology Department offers ornithological studies in the Avian Behavior Laboratory, specializing in basic and applied waterfowl research, waterfowl management techniques, and avian attachment behavior, or imprinting.

Facilities.--The Avian Behavior Laboratory provides both indoor and outdoor facilities for waterfowl. Indoor facilities are on the campus of the university in the Psychology Department. These consist of an aviary with controlled light cycles, and separate sound-deadened, heat-controlled testing chamber. The outdoor facility, at the Field Station of the Avian Behavior Laboratory maintains a flock of 75 Giant Canada Geese and about 100 Mallards as well as a few domestic mallards. Separate holding pens, boats, an observation tower, predator-control fencing, and live-in blind round out the field research facilities. Other off-campus research resources include the Delta Waterfowl Research Station, located 60 miles northwest of Winnipeg, the Northern Prairie Wildlife Research Center at Jamestown, North Dakota some 350 miles southwest of Winnipeg, Oak Hammock Marsh, 14 miles north of Winnipeg, and the Assiniboine Park-Zoo within the city.

Faculty.--Those currently involved with ornithological studies are:

  • Spencer G. Sealy, Department of Zoology: Various interests in ornithology.
  • L. James Shapiro, Department of Psychology: Director of Avian Behavior Laboratory, studies on waterfowl biology, particularly attachment behavior and its development in ducklings.
For information.--Write to Dr. L. James Shapiro, Avian Behavior Laboratory, Department of Psychology, University of Manitoba, Winnipeg, Manitoba, Canada R3T 2N2, telephone (204) 474-7244 or 9338. You may e-mail Dr. Shapiro at, and fax to (204) 474-7599



University of Michigan, Ann Arbor, MI 48109, USA

Description.-- The University of Michigan, a public university founded in 1817, enrolls approximately 35,000 students, of whom about 10,000 are graduate students. Ann Arbor, the cultural center of southern Michigan, is a small city 50 miles west of Detroit. Graduate studies of birds, leading to a degree of M.S. or Ph.D., are usually undertaken in the Museum of Zoology, Department of Biology or the School of Natural Resources and Environment. Ornithology has been a part of the university almost since its inception, especially in the Bird Division of the Museum of Zoology (see Payne, R. B. 2000. Ornithology at the University of Michigan Museum of Zoology: an historical account. In: Contributions to the History of North American Ornithology, vol. 2 (W. E. Davis and J. A. Jackson, eds). Nuttall Ornithological Society, Cambridge, MA).

Facilities.-- The Kraus Natural Science Building, Dana Building, and Ruthven Museums house student and faculty offices and laboratories. The Museum of Zoology contains an extensive ornithological library in addition to the other science libraries in the Museum and elsewhere in the University. The bird collection in the Museum is one of the largest in the world, with over 210,000 skins, skeletons, fluid-preserved specimens, eggs, and tissue samples, representing 66% of the world's bird species and 87% of bird genera. The Museum also contains the Genomic Diversity Lab, a shared-use facility for molecular studies of evolution and systematics. Aviaries are available on the roof of the Museum and indoors in the Bird Division. Several properties of the University are suitable for field work on birds. The Nichols Arboretum and Matthaei Botanical Gardens are located in Ann Arbor. The 600-ha E. S. George Reserve is 25 miles from Ann Arbor and is administered by the Museum of Zoology. The University of Michigan Biological Station, located at the northern tip of the southern peninsula of Michigan, possesses a range of habitats and facilities for ornithological field research.

Faculty.-- Many faculty members are available to direct ornithological studies. Faculty members who study birds include:

  • Bobbi Low, Professor of Natural Resources and Environment: Life history, sexual selection, and behavior, including: evolutionary and behavioral ecology of wildlife species; resource control, reproductive success, and parental strategies in vertebrates. (e-mail:
  • David Mindell, Associate Professor of Biology and Curator of Birds: Evolution and phylogeny within and among orders of birds based on DNA sequences; evolution of mitochondrial genes and gene arrangement among birds and other amniotes; assessment and application of variable rates of sequence change in molecular evolution and systematics; phylogeny and molecular evolution of retroviruses. (e-mail:
  • Robert B. Payne, Professor of Biology and Curator of Birds: Social behavior and ecology of birds and avian systematics, including: parental care, social systems, and reproductive rates in brood parasitic birds; population structure and mating systems; sexual imprinting; function, development, and evolution of vocal behavior; ecological and social adaptations of song dialects in several species of North American and African birds. (e-mail:
  • Emily Silverman, Assistant Professor of Natural Resources and Environment: behavioral and community ecology of aquatic birds; interspecific associations in mixed-species groups; probability models and variability in biological systems; nonlinear estimation; assessment and testing of dynamic models (e-mail:
For information.-- Contact one of the following offices, or the faculty member whose activities match your interests.

Department of Biology: Graduate Coordinator, Department of Biology, Natural Science Building, Ann Arbor, MI 48109-1048. (734) 764-1443. e-mail: Visit the departmental website at For more information on resources and collections in the Museum of Zoology, visit the website at

School of Natural Resources and Environment: Office of Academic Programs, School of Natural Resources and Environment, Dana Building, Ann Arbor, MI 48109. (734) 764-6453. e-mail: Visit the school website at


The University of Minnesota, St. Paul, MN 55108.

Description.--The Avian Research Center at the University of Minnesota promotes interdisciplinary research on birds. It involves nine graduate degree programs: Agricultural Engineering (M.S.Ag.E., M.Ag.E., and Ph.D.), Masters in Public Health (M.P.H.), and M.S. and Ph.D. degrees in Animal Physiology, Animal Science, Conservation Biology, Ecology, Veterinary Microbiology, Veterinary Pathology, and Wildlife Conservation. The large faculty in so many departments provide numerous and diverse opportunities for ornithological research. In addition, students may earn the M.S. and Ph.D. degrees in Zoology in the Department of Ecology, Evolution and Behavior.

Facilities.--Support facilities for avian studies at the University of Minnesota include these: Gabbert Raptor Center, specializing in the treatment and rehabilitation of birds of prey; Isolation Facilities in an experimental animal housing complex; James Ford Bell Museum of Natural History, housing extensive research collections of birds and other vertebrates; Minnesota Cooperative Fish and Wildlife Research Unit, promoting cooperative research among state natural resource agencies, the U. S. Biological Service, and the University; Natural Resources Research Institute, fostering economic development of Minnesota's natural resources in an environmentally sound manner; Poultry Teaching and Research Unit; Rosemount Turkey Research Unit; Veterinary Diagnostic Laboratory; and Wildlife Rehabilitation Clinic.

Faculty.--Those associated with avian studies are:

  • David E. Andersen, College of Natural Resources: Conservation and management of wild birds.
  • Martin E. Bergeland, Department of Veterinary Diagnostic Medicine: Animal health diagnostic service for poultry industries, companion birds, and wild birds.
  • Chuck Clanton, Department of Agricultural Engineering: Turkey waste management.
  • Craig Coon, Department of Animal Science: broiler and layer nutrition and management.
  • James Cooper, Department of Fisheries and Wildlife: Ecology and management of various species of North American waterfowl.
  • Kendall W. Corbin, Department of Ecology, Evolution and Behavior: Evolutionary ecology of the genus Zonotrichia; genetic structure of avian populations; speciation and formation of allospecies; biochemical systematics.
  • Francesca Cuthbert, Department of Fisheries and Wildlife: Conservation and biology of small avian populations.
  • Gary E. Duke, Department of Veterinary PathoBiology: Digestive physiology of poultry and raptors.
  • Mohammed El Halawani, Department of Animal Science: Neuroendocrinology of avian reproduction.
  • Douglas N. Foster, Department of Animal Science: Molecular biology of avian species.
  • James R. Kitts, Department of Fisheries and Wildlife, College of Natural Resources: Habitat management for birds.
  • D. Frank McKinney, Department of Ecology, Evolution and Behavior: Comparative ethology; evolution of social behavior and communication in birds.
  • Daniel P. Shaw, Department of Veterinary Diagnostic Medicine: Diagnostic services for poultry industry, companion birds, and wild birds.
  • Robert N. Shoffner, Department of Animal Sciences (Professor Emeritus): Genetics of poultry.
  • V. Sivanandan, Department of Veterinary PathoBiology: Control of viral respiratory diseases in poultry.
  • Paul E. Waibel, Department of Animal Sciences: Nutrition in turkeys.
  • Mary Walser, Department of Veterinary PathoBiology: Pathogenesis of noninfectious skeletal diseases in poultry.
  • Robert M. Zink, Department of Ecology, Evolution and Behavior: Ornithology, systematics.
For information.--Concerning degrees associated with the Avian Research Center, contact: Avian Research Center, University of Minnesota, 301 Veterinary Science Building, 1971 Commonwealth, St. Paul, MN 55108, telephone (612) 625-1248. Concerning the program in Zoology, contact: The Graduate Secretary, Department of Ecology, Evolution and Behavior, 100 Ecology Building, 1987 Upper Buford Circle, St. Paul, MN 55108, telephone (612) 624-6770.



University of Nebraska, Lincoln, Nebraska 68583/68588.

Description.--The University of Nebraska offers Ph.D. and M.S. degrees through two departments for students interested in ornithology: the School of Biological Sciences (BIOS) and the School of Natural Resources (SNR). The School of Biological Sciences has faculty members with interests in ecology and organismal biology, as well as cellular and molecular biology. The School of Natural Resources has faculty with interests in wildlife and forest ecology, as well as agriculture meteorology, soil and water science, and geographic information technology. Ecologists from the two schools collaborate together through the Initiative in Ecological and Evolutionary Analysis (

Facilities -- Graduate students at UNL have access to at least 26 field sites through the state of Nebraska, including forest and grassland areas near the Platte River, the sandhills, and the Rainwater Basin ( Cedar Point Biological Field Station ( is a unique facility with a major emphasis being the study of natural systems located in diverse and scenic habitats&endash;the station is at the juncture of four major grassland systems: short grass prairie, sand/sage prairie, mixed grass prairie, and sand hills. The University of Nebraska State Museum's zoological collection contains thousands of mammals, birds, reptiles, amphibians, fishes and invertebrates for research use by staff and graduate students. UNL is also the home of the Center for Advanced Land Management Information Technologies, specializing in remote sensing and Nebraska databases (

Faculty.--Those supervising studies of avian biology:



University of South Dakota, Vermillion, South Dakota 57069-2390.

Description.--The Department of biology offers the degrees of Ph.D., Master of Arts (M.A. with thesis (Plan A), and M.A. without thesis (Plan B, for teachers and other students requiring broad training in various aspects of biology). Teaching and Research Assistantships are available for graduate students. The Department of Biology has 13 faculty members with research programs that include the areas of conservation ecology and genetics, community ecology, ecological physiology, endocrinology and neuroendocrinology, developmental biology and molecular biology. A major research emphasis of the department is in the area of stress biology, investigating how animals and plants respond to stressful environments or stressful situations.

Facilities.--The department has a modern computer laboratory for graduate students, aquarium and greenhouse facilities, and an herbarium. The W. H. Over Museum of Natural and Cultural History is also located on campus and houses a large collection of bird skins, specializing in birds of the northern Great Plains region..

Faculty.--Those supervising studies of avian biology:

  • Hugh B. Britten: conservation genetics, conservation biology, endangered species.
  • David L. Swanson: ecology and energetics of migrating passerines, cold tolerance of small birds.
For information.--Write to Graduate Steering Committee, Department of Biology, University of South Dakota, Vermillion, SD 57069-2390, (605) 677-6175.


University of Wisconsin-Green Bay, Green Bay, WI 54311-7001, USA.

Description.--The University of Wisconsin-Green Bay offers M.S. degrees in Environmental Science and Policy with an emphasis in Ecology and Biological Resources Management. This program includes an active program of ornithological research with studies of bird metapopulation dynamics, wetland birds, forest management and bird populations, GIS applications in ornithological research, and applied avian ecology. UWGB is a regional institution of approximately 5000 students, located on the shores of Green Bay. Its programs emphasize interdisciplinary, problem-focused studies, with emphasis on the environment.

Facilities.--The Richter Museum of Natural History, part of the Department of Natural and Applied Sciences, holds one of North America's largest collections of bird eggs.

Faculty.--Those involved in ornithological studies:

  • Gregory J. Davis: mathematical modeling of metapopulations; landscape analysis of bird populations; smooth, discrete, and chaotic dynamical systems.
  • Hallet J. Harris: animal and wetland ecology; wildlife management; ecosystems management.
  • Robert W. Howe: terrestrial ecology and conservation biology; bird population dynamics: habitat fragmentation and landscape ecology.
  • Thomas C. Erdman, Associate Curator, Richter Museum of Natural History (staff): Raptor biology; natural history of birds.
For information.--Contact the Office of Graduate Studies, TH 335, University of Wisconsin-Green Bay, 2420 Nicolet Drive, Green Bay, WI 54311-7001



University of Arkansas, Fayetteville, AR 72701, USA.

Description.--The University of Arkansas, Fayetteville, located in the scenic Ozark Plateau region of northwestern Arkansas, has had an active graduate program in ornithology since the late 1950s involving 10 to 20 graduate students in a given year. Students may earn either M.S. or Ph.D. research degrees in the Department of Biological Sciences. The emphasis is on avian ecology and behavior, endangered species, and tropical ornithology.

Facilities.--Extensive forests and a diversity of associated habitats exist near the University in the Ozark National Forest, Ouachita National forest, Buffalo National River and Pea Ridge National Military Park where research projects have been conducted. The fine facilities of the Ozark Natural Science Center are available in the Madison County Wildlife Management Area, as is the Lake Fayetteville Environmental Center close to town. There are several large reservoirs in the area for water bird studies. In addition, students have been doing research in Belize, Costa Rica and India. Laboratory facilities and equipment include controlled environmental chambers and telemetry apparatus for conducting laboratory and field studies requiring the measurement of metabolic rates, and movements. Cooperative relations exist with the Department of Poultry Science, College of Agriculture, provides expertise in avian physiology and access to the analytical avian nutrition facilities. The Center for Advanced Spacial Technologies provides Geographic Information Systems (GIS) and related technologies that are widely available to researchers. Opportunities exist with the Arkansas Cooperative Fish and Wildlife Research Unit, U.S. Geological Survey, which is a functional entity within the department.

Faculty.--Those involved in ornithological studies:

  • Douglas A. James, Department of Biological Sciences.
  • David G. Krementz,
  • William L. Thompson,
  • Kimberly G. Smith, Department of Biological Sciences.
For information.--Write to Department of Biological Sciences, University of Arkansas, Fayetteville, AR 72701, and request materials and applications for graduate studies including the document that summarizes the research activities of the faculty. It is also important to make contact with and gain support from a specific faculty member. More information concerning the departmental program and faculty interests is available on the departmental web site:

Arkansas State University, State University, AR 72467, USA.

Description.--Arkansas State University is located on the Mississippi Delta in the city of Jonesboro approximately 70 miles northwest of Memphis, TN. Recent emphasis of the graduate program in Biology includes studies in Conservation Biology, Avian Ecology, Ecotoxicology, Wildlife Management, and other environmental and applied ecological sciences. Students may earn an M.S. degree.

Facilities.--The Department of Biological Sciences occupies approximately 60,000 square feet of offices and research laboratories within three buildings. A modern Remote Sensing and GIS facility (900 ft2) and an Ecotoxicology Research Laboratory (3500 ft2) are available for use by ornithology students and faculty. Most ornithology students have offices within or have access to the Population Ecology Laboratory (680 ft2). A variety of standard field equipment is available for ornithological studies (such as mist nets, tape recorders, radio telemetry instruments, spotting scopes, etc.). The faculty maintains active relationships with local federal and state land management agencies, and conducts field research at nearby national wildlife refuges, national forest and state wildlife management areas.

Faculty.--Those currently involved in ornithological studies are:

  • James Bednarz, Department of Biological Sciences:
  • Rick McDaniel, Department of Biological Sciences:
For information.--Dr. James Bednarz, Department of Biological Sciences, Arkansas State University, P. O. Box 599, State University, AR 72467


Friends University, Wichita, KS 67213, USA.

Description.--Friends University offers M.S. degrees in Environmental Studies. Students may choose a concentration area that exposes them to various areas of avian conservation and management.

Facilities.--Students may work with the Curator of Birds at the Sedgwick County Zoo on a Practicum and research project. Students interested in other aspects of birds may work with a faculty member or with a bird biologist associated with a state agency.

Faculty.--The faculty member currently involved in ornithological studies is:

  • Alan D. Maccarone, Biology Department and Director of Environmental Studies.
For information.--Dr. Alan D. Maccarone, Biology Department, 2100 University, Wichita, KS 67213.


Kansas State University, Manhattan, KS 66506, USA.

    Description.&emdash;Kansas State University was founded in 1863 as a land-grant research institution. The 269 ha campus is located 200 km west of Kansas City in the rolling Flint Hills of northeast Kansas. Current enrolment is about 20,000 students. The Division of Biology offers graduate programs leading to Master of Science and Ph.D. degrees. Kansas State has a particularly strong research program on the ecology of grassland ecosystems. Individual faculty have expertise in quantitative population statistics, demographic modeling, landscape and conservation ecology, and the ecology of tropical and grassland birds. Graduate students are supported with research and teaching assistantships. Financial support is guaranteed while progress is being made towards completion of a degree. Manhattan is a small college town with good services and a low cost of living.

    Facilities.&emdash;Kansas State University offers an excellent range of facilities to support graduate research on birds. Konza Prairie Biological Station is a 3,487 ha native tallgrass prairie preserve and is one of a network of 24 Long-Term Ecological Research Sites (LTER) funded by the National Science Foundation. Long-term research has addressed the effects of different grazing and fire treatments on population and community dynamics of the flora and fauna of tallgrass prairie. The station is a 20 min drive south of the university and supports a diverse avifauna. Research facilities at Konza Prairie include: lab space, a wood and metal shop, and project vehicles. Campus facilities of the Division of Biology include: computer labs (including GIS), greenhouses, good library holdings, a transmission electron microscope, and a stable isotope mass spectrometry laboratory. Museum holdings of avian specimens and facilities for animal-care are somewhat limited. The Kansas Cooperative Fish and Wildlife Unit has an office on campus and is a joint cooperative unit between USGS-BRD, the Kansas Dept. of Wildlife and Parks and the university.

    Faculty.&emdash;Faculty members currently involved in ornithological studies include:

  • C. Dusty Becker, Department of Horticulture, Forestry and Recreation Resources: Community ecology of tropical birds, effects of recreational use on riparian birds.Ted T. Cable, Department of Horticulture, Forestry and Recreation Resources: Human dimensions of wildlife management.
  • Jack F. Cully, Jr., Division of Biology: Habitat relationships and conservation of grassland birds, brood parasitism, and wildlife diseases.
  • Philip S. Gipson, Division of Biology: Responses of bird communities to environmental disturbance and habitat enhancement.
  • David A. Rintoul, Division of Biology: Lipid biochemistry, physiology of avian migration.
  • Robert J. Robel, Division of Biology: Wildlife ecology, population dynamics of upland gamebirds.
  • Brett K. Sandercock, Division of Biology: Demography, population biology and behavioural ecology of grassland and tropical birds.
  • Fred E. Wilson, Division of Biology: Reproductive physiology, hormonal control of timing of breeding.
  • Kimberly A. With, Division of Biology: Landscape and conservation ecology, effects of habitat loss and fragmentation on extinction risk.

    For information.&emdash;Write to the Graduate Selection Committee, Division of Biology, Kansas State University, 232 Ackert Hall, Manhattan, KS 66506-4901. Phone: (785) 532-6615, Fax: 785-532-6653, Home Page:



Louisiana State University, Baton Rouge, LA 70803-3216, USA.

Description.--Louisiana State University is a land-grant and sea-grant research institution located on the east bank of the Mississippi River in the capital of Louisiana. Students may earn both M.S. and Ph.D. degrees in the Department of Biological Sciences and in the School of Forestry, Wildlife, and Fisheries. Students may receive full assistantships for either 12 or 9 months; duties involve either teaching laboratory sections for courses offered by the Department of Zoology and Physiology or curating in the bird or frozen tissue collection of the Museum of Natural Science.

Facilities.--LSU offers a strong program in systematics and evolutionary biology. The LSU Museum of Natural Science (LSUMNS) has an extensive collection of neotropical birds (170,000 specimens) and frozen tissues of birds (3,000 species represented) and a strong history of ornithological research since its founding in 1936. The LSUMNS program in ornithology has three distinct but interrelated areas of graduate training: fieldwork in the tropics, recently concentrating on Peru and Bolivia; specimen-based research using museum collections; and biochemical systematics. Several faculty members in the School of Forestry, Wildlife, and Fisheries also are involved in ornithological research, with strong emphasis on waterfowl and shorebird ecology.

Faculty.--Those currently involved in ornithological studies are:

  • Dominique Homberger, Department of Biological Sciences: Functional and evolutionary morphology of birds.
  • J.V. Remsen, Museum of Natural Science (MNS is affiliated with Department of Biological Sciences): Ecology, zoogeography, and geographic variation of tropical birds.
  • Fred H. Sheldon, Museum of Natural Science (MNS is affiliated with Department of Biological Sciences): Molecular systematics of birds; ecophylogenetics; biogeography of birds of Borneo.
  • Alan D. Afton, School of Forestry, Wildlife, and Fisheries: Bioenergetics of birds, especially of waterfowl.
  • Michael Chamberlain, School of Forestry, Wildlife, and Fisheries: Upland gamebird ecology.
  • Robert J. Hamilton, School of Forestry, Wildlife, and Fisheries: Feeding behavior and population ecology of birds, especially Neotropical migrants.
  • Frank C. Rohwer, School of Forestry, Wildlife, and Fisheries: Ecology of birds, especially of waterfowl.
For information.--Contact the respective department:

Department of Biological Sciences: (225) 578-1132.

School of Forestry, Wildlife, and Fisheries: (225) 578-4131.


Southeastern Louisiana University, Hammond, LA 70402, USA.

Description.--Southeastern Louisiana University is located approximately one hour from both New Orleans and Baton Rouge in a city with a population of about 20,000. At SELU, students may earn the M.S. degree. Most graduate students do field research. Teaching assistantships (with complete tuition waiver) are available to qualified students.

Facilities.--Biology facilities include the Turtle Cove Biological Research Station, approximately 20 miles south of SELU, as well as equipment such as canoes, powerboats, and trucks needed to do local field work. SELU has a teaching collection of bird skins.

Faculty.--A faculty member currently involved in ornithological studies is:

  • Philip C. Stouffer, Department of Biological Sciences: Conservation of birds; tropical ecology; effects of landscape alteration on tropical and temperate birds
For information.--Contact Dr. Philip C. Stouffer, Graduate Coordinator, Dept. of Biological Sciences, Southeastern Louisiana University, Hammond, LA 70402-0736 (504) 549-2191. E-mail:


Tulane University, New Orleans, LA 70118, USA.

Description.--The Department of Ecology, Evolution, and Organismal Biology offers M.S. and Ph.D. degree programs. All graduate students are supported by either teaching assistantships or research assistantships or, in a few cases, four-year fellowships.

Facilities.--Proximity to a number of wetland areas of Louisiana, including Bayou Sauvage National Wildlife Refuge and Jean LaFitte National Historical Park, where Tulane students work on a variety of projects. Institutional links to Dartmouth College and Hubbard Brook Experimental Forest, NH, where Tulane students work on research projects, and University of West Indies, where Tulane students carry out ongoing research in Jamaica on wintering ecology of Neotropical migrant birds.

Faculty.--A faculty member currently involved in ornithological studies is:

  • Thomas Sherry, Department of Ecology, Evolution, and Organismal Biology: Population ecology of Neotropical-Nearctic migratory landbirds; population and toxic waste monitoring in colonial wading birds; and feeding and dietary ecology in birds. (

For information.--Graduate Director, Department of Ecology, Evolution, and Organismal Biology, Tulane University, New Orleans, LA 70118.



Oklahoma State University, Stillwater, OK 74078, USA .
Description .- The Oklahoma State University in Stillwater is located in the Osage Plains roughly midway between Tulsa and Oklahoma City. Serious ornithological study in Oklahoma has its roots in Margaret Morse Nice's Birds of Oklahoma (1924), and includes outstanding contributions from other notables such as Frederick and Marguerite Baumgartner and George Miksch Sutton. Graduate students have pursued ornithological study in OSU's Department of Zoology ( since at least 1948, when the Department became the home base for Oklahoma's Cooperative Fish and Wildlife Research Unit. The Department currently offers M.S. and Ph.D. degrees in Zoology and Wildlife and Fisheries Ecology with several faculty engaged in field ornithology. Additional faculty in the College of Agricultural Sciences and Natural Resources (especially Rangeland Ecology and Management and Forestry) are also involved in avian research. Topics of research tend to emphasize ecology, conservation, and management of native birds.

Facilities .-The Department of Zoology occupies a six-floor building with offices, classrooms, laboratories, and animal rooms. A broad range of instrumentation is available for research as well as teaching. The Department of Zoology also houses the OSU Vertebrate Research Collections. Available for use in field studies is the University-owned Lake Carl Blackwell area, the Cross Timbers Experimental Range, and access to multiple public lands across the state managed by cooperators and partners.

Faculty .--Those involved primarily in ornithological studies:
Craig A. Davis, Dept. of Zoology. (
Timothy J. O'Connell, Dept. of Zoology. (
For information .- For general inquiries regarding graduate study in ornithology at OSU, please contact Tim O'Connell. Specific advice for applicants may be found here:



Stephen F. Austin State University, Nacogdoches, TX 75962, USA.

Description.--Stephen F. Austin State University is a state-funded university located in the "pineywoods" of eastern Texas; Nacogdoches has a population of about 30,000 and is located 165 miles north of Houston. The Department of Biology offers an M.S. degree with emphasis in ornithology. Students may receive teaching assistantships.

Facilities.--The Department of Biology works closely with biologists, including several ornithologists, at the U.S. Forest Service Research Station in Nacogdoches.

Faculty.--A faculty member currently involved in ornithological studies is:

  • Cliff Shackelford, Department of Biology: Ecology and behavior of birds.

    For information.--Contact Dr. Don Hay, Chair, Department of Biology, Box 13003, Stephen F. Austin State University, Nacogdoches, TX 75962, (409) 468-3601.


The Arthur Temple College of Forestry at Stephen F. Austin State University

Description.--Accredited by the Society of American Foresters, the Arthur Temple College of Forestry (College) is dedicated to meeting the teaching, research, and service needs of professional managers of land and related forest resources. Its primary purpose is to provide students with the education and field experience required for success beyond the campus. A secondary purpose is to provide the knowledge and expertise required when solving problems of resource management and use. This includes research and continuing education as well as graduate education and technology transfer. College ( faculty build on strong working relationships with resource managers for exemplary programs in teaching, research, and service.

Faculty.&emdash;A faculty member currently involved in ornithological studies is:

  • Warren C. Conway, Assistant Professor of Wildlife Management, phone: 936-468-2090, email:

    For information.&emdash;Director of Graduate Studies in Forestry, Stephen F. Austin State University, Arthur Temple College of Forestry, Box 6109, Nacogdoches, TX 75962 Telephone: (936) 468-3301


Texas Tech University, Lubbock, Texas 79409-3131.

Description.--The Department of Biological Sciences is large enough to meet the research and instructional needs of a diversity of students, including those interested in ornithological studies. The focus is on Environmental Sciences and Ecology.

Facilities.--The Institute for Environmental Studies provides a broad spectrum of support for graduate students in ecology and environmental sciences, ranging from research fellowships to an array of 75 scientists in other departments and colleges with whom a student may interact. The natural Sciences Research laboratory (The Museum), includes a collection of 4500 bird specimens, and a cryogenic tissue collection; the International Center for Arid and Semiarid Land Studies, Fish and Wildlife Cooperative Unit, and The Junction Center biological field station support graduate student research projects.

Faculty.--Faculty members currently involved with studies adaptable to ornithological problems are:

  • M. Kent Rylander, Ornithology and Comparative Neuroanatomy, social behavior of geese, role of birds in modified desert ecosystems.
  • Michael R. Willig, Population and Community Ecology.
For information.-Dr. Michael R. Willig, Chair, Ecology Program, Department of Biological Sciences, Texas Tech University, Lubbock, TX 79409-3131


The University of Texas - Pan American, Edinburg, TX 78539-2999, USA.

Description.--The University of Texas-Pan American is located in the Lower Rio Grande Valley, and a variety of temperate and tropical habitats are available within a day's drive. Several subtropical habitats are available in the Lower Rio Grande Valley, including Tamaulipan thorn forest, Rio Grande floodplain forests, grasslands, agricultural habitats, and coastal lagoons and mudflats. The Department of Biology offers an M.S. degree, in which it is possible to specialize in ornithology. Most graduate students are supported with teaching assistantships, and some graduate students receive research assistantships.

Facilities.--The Coastal Studies Laboratory at South Padre Island provides facilities for research. Vehicles and boats for field research are available.

Faculty.--A faculty member currently involved in ornithological studies is:

  • Tim Brush, Department of Biology: Habitat use and population ecology of subtropical birds, with special interest in conservation biology.
For information.--Contact: Director of Graduate Studies in Biology, Department of Biology, The University of Texas-Pan American, Edinburg, TX 78539 Telephone: (956) 387-3537




Simon Fraser University, Burnaby, BC, Canada.

Description.--SFU Biosciences offers MSc and PhD degrees in behavioral ecology, physiolgical ecology (reproduction and migration) and population biology, with emphasis on migratory waterfowl, seabirds, and shorebirds. Qualified students may obtain Graduate fellowships.

Facilities.--Permanent research facilities are maintained on Triangle Island (BC’s largest seabird colony), at Riske Creek (wetland/grassland/forest ecosystem) and at Creston (interior wetland). Other research projects are based in Desolation Sound (Marbled Murrelets) and the Fraser Estuary. Laboratory facilities are available, including a range of physiological/endocrinological methods.

Faculty.--Faculty currently involved in ornithological studies are:

  • Dr. Evan Cooch: population dynamics, demography, life history theory, and applied statistics.
  • Dr. Fred Cooke: population biology and evolutionary ecology of birds.
  • Dr. David B. Lank: behavioral ecology, alternative mating strategies, and behavioral genetics.
  • Dr. Tony D. Williams: reproduction and migration physiology.
  • Dr. Ron Ydenberg: evolutionary ecology; foraging and social behavior, predator-prey interactions.
For information.--Contact: Mrs. Sylvia Foran, Graduate Secretary, Biosciences, Simon Fraser University, 8888 University Drive, Burnaby, BC, V5A 1S6, Canada, or contact the individual faculty members listed above.


Idaho State University, Pocatello, ID 83209, USA

Description.--Idaho State University is located at the edge of the Snake River plain in SE Idaho, along the Old Oregon Trail at the confluence of the Portneuf and Snake Rivers. ISU offers both M.S. and Ph.D. degrees in the Department of Biological Sciences, with a strong program in Ecology. Qualified students may receive graduate or research assistantships.

Facilities.--The Idaho Museum of Natural History, located on the ISU campus, houses a large collection of vertebrates and one of the largest collection of vertebrate fossils in the country. ISU has a field research station at Harriman State Park, 30 miles south of Yellowstone National Park, and the Department of Biological Sciences has the O'Neal Ecological Reserve, 200 acres located about 25 miles south of Pocatello. The Center for Ecological Research and Education (CERE) was formed in 1989 to serve as a focus for the strong ecology program at ISU; CERE was formed to facilitate individual research in Ecology and to provide a framework for collaborate interdisciplinary teaching and research based on ecological principles and problems.

Faculty.--The faculty member currently involved in ornithological studies is:

  • Charles H. Trost, Department of Biological Sciences: Status and distribution of birds in Idaho; reproductive success and social behavior of birds.
For information.--Dr. Rod Seely, Department of Biological Sciences, Campus Box 8007, Idaho State University, Pocatello, ID 83209-8007.


Oregon State University, Corvallis, OR 97331

Description:-- Oregon State University ( is a land grant institution established in 1848 in Corvallis, Oregon (population ~ 50,000). Located approximately 80 miles south of Portland, 50 miles east of the Pacific Ocean, and 60 miles from the Cascade mountains, OSU is perfectly situated for field and laboratory investigations in many ecosystems and on innumerable topics in ornithology. The departments listed below offer M.S. and Ph.D. degrees. In addition, graduate students in ornithology benefit from the numerous agency cooperators on campus. Thus, graduate faculty members can include employees of the U.S. Geological Survey, Environmental Protection Agency, U.S. Forest Service, National Oceanic and Atmospheric Administration, etc.

Facilities:--Oregon State University's Hatfield Marine Science Center ( is a research and teaching facility located in Newport, Oregon on the Yaquina Bay estuary, about one mile from the open waters of the Pacific Ocean and 50 miles from Corvallis. HMSC plays an integral role in programs of marine and estuarine research and instruction and as a laboratory facility serving resident scientists as a base for far-ranging oceanographic studies.

The Center for Gene Research and Biotechnology ( serves the biosciences research community at Oregon State University with the ultimate goals of improving health, sustaining natural and agricultural resources, and preserving environmental quality. The Center offers leadership and services to faculty, staff and students through core facilities, seminars, and retreats. It also provides a focal point for researchers to make contacts, initiate collaborations, and establish new technologies in their own laboratories.

Nestled in the central Cascade Range of Oregon, the H.J. Andrews Experimental Forest ( is a living laboratory that provides unparalleled opportunities for the study of forest and stream ecosystems. During the last 21 years as a part of the National Science Foundation Long Term Ecological Research (NSF-LTER) program, the Andrews Experimental Forest has become a leader in the analysis of forest and stream ecosystem dynamics. Long-term field experiments and measurement programs have focused on climate dynamics, streamflow, water quality, and vegetation succession. Currently researchers are working to develop concepts and tools needed to predict effects of natural disturbance, land use, and climate change on ecosystem structure, function, and species composition.

USGS Forest and Rangeland Ecosystem Science Center ( FRESC combines research scientists and support staff originating from the Bureau of Land Management, National Park Service, and U.S. Fish and Wildlife Service in Oregon, Washington, and Idaho. The result is an integrated facility focused on providing biological information to land managers in the Department of the Interior and others with natural resource concerns in the western U.S. and adjacent states and Canadian provinces.

USGS Oregon Cooperative Fish and Wildlife Research Unit ( OR_FW): The Wildlife Program of the Oregon Coop. Unit trains graduate students and conducts wildlife research of interest to our state and federal cooperators as well as other natural resource management agencies/groups using students, faculty, research assistants, and facilities of Oregon State University. The Unit interprets and disseminates research results to the scientific community, natural resource agencies, and the public and provides technical assistance to natural resource managers. Current emphasis in the Unit focuses on Pacific Northwest forest-wildlife management issues and migratory waterbirds.

Faculty Associated with Ornithology:--

  • Robert G. Anthony (Department of Fisheries and Wildlife, USGS Oregon Coop. Unit): population biology/demography, particularly of raptors such as Spotted Owls; contaminants.
  • Bruce G. Dugger (Department of Fisheries and Wildlife): Ecology and conservation of waterbirds, particularly waterfowl; restoration and management of wetlands; behavioral ecology.
  • Katie M. Dugger (Department of Fisheries and Wildlife): Avian population ecology particularly the effects of climate and global climate change on survival and reproductive, foraging ecology, waterbird management and conservation
  • W. Daniel Edge (Department of Fisheries and Wildlife): avian habitat relationships.
  • Paul Farber (Departments of History and Zoology): history of ornithology and natural history -- bird-watcher watcher.
  • Eric D. Forsman (Department of Fisheries and Wildlife, U.S. Forest Service Pacific Northwest Forest Experiment Station): Ecology of forest birds and mammals, spotted owls, pygmy owls, barred owls, red tree voles, genetics, dispersal, population analysis, home range, habitat selection, diets, forest management.
  • Susan M. Haig (Department of Fisheries and Wildlife, USGS FRESC): conservation genetics, behavioral ecology including mating systems and dispersal patterns, shorebird/wetland landscape conservation, conservation of endangered Micronesian avifauna.
  • John P. Hayes (Forest Science Department): habitat ecology, influences of natural disturbance and forest management on songbirds.
  • Charles J. Henny (Department of Fisheries and Wildlife, USGS FRESC): environmental contaminants, long-term biomonitoring, fish-eating birds and raptors.
  • Patricia L. Kennedy (Department of Fisheries and Wildlife): ecology and management of forest and grassland avifauna; regulation of avian populations; raptor ecology and management; effects of livestock grazing on avian populations and communities.
  • Arch McCallum (Applied Bioacoustics, Inc.): Bioacoustics, particularly conservation applications. Conservation biology, especially theory and methodology of monitoring and habitat assessment. Flammulated Owls. Evolution of the Paridaes.
  • David K. Mellinger (Hatfield Marine Science Center) acoustic monitoring of wildlife populations; nocturnal flight calls of migrating birds; signal processing methods for automatic recognition of animal sounds.
  • Laura Nagy (Department of Fisheries and Wildlife, Environmental Protection Agency): Population modeling, behavioral ecology, risk assessment.
  • Fred Ramsey (Department of Statistics): Statistical analyses, author of Field Guide to Birds of Oregon.
  • W. Douglas Robinson (Department of Fisheries and Wildlife): avian conservation, aridlands ecology, tropical ecology.
  • Tara Robinson (Department of Fisheries and Wildlife): avian ecology, behavioral ecology, tropical ecology, genetics.
  • Daniel Roby (Department of Fisheries and Wildlife, USGS Oregon Coop. Unit): Physiological ecology, reproductive energetics, seabird/fisheries interactions, ecology of colonial waterbirds, restoration following oil spills.
  • Richard Schmitz (Department of Fisheries and Wildlife): spatial ecology, upland game birds.
For Information:--

Graduate Admissions: 541-737-4881,

Fisheries and Wildlife Department: 541-737-4531,

Forest Science Department: 541-737-2244,

History Department: 541-737-3421,

Statistics Department: 541-737-3366,

Zoology Department: 541-737-3705,


Portland State University, Portland, OR 97207, USA

Faculty.--The faculty member currently involved in ornithological studies is:
  • Dr. Michael T. Murphy, Department of Organismal Biology: Population ecology, habitat use, avian behavior, and the ecology of migrants in the Caribbean.
For Information.--Dr. Michael Murphy, Department of Organismal Biology, P.O. Box 751, Portland State University, Portland, OR 97207-0751



University of Washington, Seattle, WA 98195, USA.

Description.--The University of Washington is located within easy access to many different environments, including marine waters and estuaries, temperate rain forest of the Olympic Peninsula, the Cascade Mountains, and shrub-steppe of eastern Washington. The University is well-located for studies of avian ecology; during winter, there are large concentrations of waterfowl and raptors on or near campus. Students may earn M.S. and Ph.D. degrees in the Department of Zoology and in the Wildlife Science program in the College of Forest Resources. All Ph.D. students in the Department of Zoology are guaranteed six years of support, either through teaching or research assistantships or through fellowships or traineeships; all students in the Wildlife Science program are supported by research assistantships.

Facilities.--The Burke Museum contains a research collection of about 40,000 bird specimens and a new collection of frozen tissues, and supports student research done in the field or on preserved specimens. The Friday Harbor Laboratories maintain a close affiliation with the Department of Zoology. The Wildlife Science program has an avian ecology graduate program, in collaboration with the Washington Cooperative Fish & Wildlife Unit and the Washington Dept. of Fish and Wildlife.

Faculty.--Faculty currently involved in ornithological studies are:

  • Dee Boersma, Department of Zoology: Conservation of birds; seabirds.
  • Scott Edwards, Department of Zoology: Molecular systematics and evolution of birds.
  • Gordon Orians, Department of Zoology: Ecology of birds; icterid social systems.
  • Sievert Rohwer, Department of Zoology: Evolution of birds; molts and coloration.
  • Michael Beecher, Department of Psychology: Mechanisms, development, and function of bird song.
  • Eliot Brenowitz, Department of Psychology: Integration between mechanism and function in animal behavior, with emphasis on acoustic communication in birds.
  • John Wingfield, Department of Zoology: Environmental endocrinology of birds.
  • David Manuwal, Wildlife Science Group, College of Forest Resources:
  • Christian Grue, Leader, Washington Cooperative Fish & Wildlife Research Unit:
  • Stephen Penland (Affiliate faculty), Washington Dept. of Fish and Wildlife:
For information.--Graduate Program Coordinator, Department of Zoology, NJ-15, University of Washington, Seattle, WA 98195 (206-685-8240), or David A. Manuwal, Wildlife Science Group, College of Forest Resources, AR-10, University of Washington, Seattle, WA 98195 (206/543-1585).



University of Arizona, Tucson, AZ 85721, USA.

Description.--The Department of Ecology and Evolutionary Biology at the University of Arizona emphasizes doctoral education, and the M.S. degree is also available. One of the goals of the Department of EEB is to develop studies in avian biology with members of the birding community in nearby areas, such as Ramsey and Madera Canyons, Portal, and Patagonia; such areas already provide extensive research opportunities for faculty and students. The Department of EEB normally guarantees financial support to successful Ph.D. candidates for five years; M.S. students are not guaranteed financial support. Financial support for doctoral research and attendance at professional meetings is available from the Department, the University, and through a Research Training Grant in Biological Diversification.

Facilities.--The University of Arizona sponsors more than 60 cooperative Facilities and Services, including the Laboratory of Molecular Systematics and Evolution, Arizona Fish and Wildlife Cooperative Research Unit, Environmental Research Laboratory, Arizona-Sonora Desert Museum, and Southwest Research Station. The Department of EEB has a bird collection of 14,000 catalogued study skins, 3,000 skeletons, and wings and tails from 800 specimens, and access to the Santa Rita Experimental Station, 50,000 acres of shrub-grassland and oak woodlands, approximately 30 miles south of Tucson.

Faculty.--Faculty currently involved in ornithological studies are:

  • William A. Calder, Department of Ecology and Evolutionary Biology: Physiological ecology of birds, biological scaling; conservation biology.
  • Steven Hopp, Adjunct in Department of Ecology and Evolutionary Biology: Comparative biology, particularly vocal behavior, of vireo species; acoustic analysis.
  • Tom Huels, Department of Ecology and Evolutionary Biology : Curatorial specialist in charge of collections.
  • Irene M. Pepperberg, Department of Ecology and Evolutionary Biology: Cognitive and communicative behavior of parrots; effects of social interaction or vocal learning in birds.
  • Stephen M . Russell, Department of Ecology and Evolutionary Biology (emeritus): Migration patterns of hummingbirds.
For information.--Contact: Chair, EEB Graduate Committee, Biological Sciences West Building, Room 310, University of Arizona, Tucson, AZ 85721 (602/621-1165).



University of California, One Shields Avenue, Davis, CA 95616, USA.

Description.--Davis is a residential community of about 60,000 in the Sacramento Valley, 15 miles west of Sacramento and 72 miles northeast of San Francisco. UCD is particularly strong in the biological sciences, with over 50 biological science departments, divisions, and graduate groups on campus, in addition to the veterinary and medical schools. Students interested in ornithological research can pursue M.S. or Ph.D. degrees through such Graduate Groups as Ecology, Population Biology, Avian Sciences, Physiology or numerous others. The Center for Avian Biology acts as a connecting point for faculty and researchers across the Davis Campus and throughout California. Financial aid is available in the form of fellowships and teaching and research assistantships.

Facilities.--Research facilities include the University of California Natural Reserve System (NRS), which includes 27 reserves encompassing 68,000 acres and a wide range of habitats within the state. Many of these reserves have equipped field stations, with laboratory, cooking and dormitory facilities and an academic staff. The Bodega Marine Laboratory is located on Bodega Head, about 100 miles from Davis. The laboratory property fronts on both the Pacific Ocean and Bodega Harbor, and it is part of the NRS.

Faculty. Members currently involved in ornithological studies are:

  • Daniel W. Anderson, Wildlife and Fisheries Biology: Ecotoxicology and residue accumulation patterns in migratory waterbirds. Avian ecology: population, behavior, and community studies of marine birds.
  • Dale Brooks, Director, California Raptor Center: Raptor medicine and rehabilitation. Nicky Clayton, Neurobiology, Physiology and Behavior: Memory and the hippocampus in food-storing birds; brain behavior of birdsong.
  • Mary Delany, Animal Science: Embryonic development; genetics; cytogenetics.
  • John Eadie, Wildlife & Fisheries Conservation Biology: Evolution of birds; parasitism; population dynamics.
  • D. Michael Fry, Avian Sciences Department: Wildlife toxicology, environmental chemistry. Effects of toxicants on reproduction and development of birds, and the physiological and reproductive effects of oil pollution on seabirds.
  • Michael L. Johnson, Civil and Environmental Engineering: Ecology and evolution of birds and mammals, especially interactions involving spatial heterogeneity, dispersal and population dynamics. Modeling, ecotoxicology, and ecosystem risk assessment.
  • Annie King, Animal Science: Prevention of lipid peroxidation; association of protein with by-products of lipid oxidation.
  • Kirk Klasing, Animal Science: Nutrition and immunity; comparative avian nutrition.
  • Linda Lowenstine, VM: Pathology, Microbiology and Immunology: pathogenesis and pathology of spontaneous diseases of captive and free-ranging wild animals, especially birds, primates, and marine mammals; aviculture embryonic pathology.
  • Peter Marler, Evolution and Ecology: Animal behavior and behavioral ecology; ethology and neurobiology of bird song and its development with special reference to sensitive periods for learniing.
  • Joy Mench, Animal Science: Social behavior; animal welfare; environmental enrichment.
  • James R. Millam, Animal Science: Physiological basis, particularly endocrine, of how environmental and social cues modulate progression of the avian sexual cycle; captive biology of parrots.
  • Gabrielle Nevitt, Neurobiology, Physiology and Behavior: Avian olfaction.
  • Pat Wakenell, VM: Population Health and Reproduction: Viral diseases of poultry; immunology of chicken, turkey; clinical pathology of birds.
  • Wesley W. Weathers, Animal Science: Wildlife ecology and conservation, with emphasis on ecological energetics and thermal relations of wild birds, physiological adaptation to environmental stress, and allometric analysis and scaling of animal function.
  • Barry Wilson, Animal Science: Muscle growth and development; neurotoxicology; ecotoxicology
For information. For more information on these or other faculty with avian interests, please contact: Center for Avian Biology, 3202 Meyer Hall, University of California, One Shields Avenue, Davis, CA 95616. Telephone: (530) 754-8560. Fax (530) 752-4508.



Humboldt State University

    Description: HSU offers 4-8 semester courses (lectures and labs) in ornithology. These include: ornithology, ornithology II, waterfowl ecology and management, shorebird ecology and management, and birds and human society (for non-science majors). We offer three other courses on an irregular basis: ecology and management of upland game birds, ecology and management of raptors, ecology and management of seabirds. The department has seven full-time faculty, three of whom are ornithologists. Other faculty, however, direct graduate students on thesis related to ornithology. We have a 12,000 specimen teaching collection housed in the Wildlife museum, which includes live mount displays in hallways and rooms of the Wildlife and Fisheries Building. The collection is curated by a half-time staff member.  

    Facilities: A game pens facility houses live waterfowl, galliforms and columbiforms for use in teaching and research. The department runs a banding station at the Wright Urban Wildlife Refuge in Eureka. Lastly, the diverse habitats of the north coast of California provide valuable opportunities for students to perfect their field skills.  


  • Jeffrey Black,
  • Mark Colwell,
  • Luke George,
  • Matt Johnson,

    For information

    Natural Resources Graduate Program
    College of Natural Resources and Sciences
    Humboldt State University
    Arcata, CA 95521
    Forestry Building, Room 101
    (707) 826-3256  



University of Nevada Las Vegas, Las Vegas, NV 89154, USA

Description.--The Department of Biology at the University of Nevada Las Vegas offers graduate programs leading to both M.S. and Ph.D. degrees. Both programs emphasize comprehensive training in biology, and both have substantial research components. The Biology Department generally offers two years of support for qualified M.S. students, and four years for qualified Ph.D. students.

Facilities.-- The primary facilities for ornithological research are contained in the Barrick Museum of Natural History. Barrick Museum facilities consist of a rapidly growing collection consists of 7,500 research skins and 3,000 tissues, as well as a self-contained DNA laboratory.

Faculty.-- Faculty currently involved in ornithological studies are:

  • Shawn Gerstenberger, Department of Environmental Studies: Avian toxicology.
  • John Klicka, Barrick Museum, Adjunct in Department of Biology: Molecular systematics and evolution of birds; Distributions of Honduran birds.
  • Gary Voelker, Barrick Museum, Adjunct in Department of Biology: Molecular systematics and evolution of birds; molts and plumages; Distributions of Honduran birds.
For information.-- Contact: (Barrick Museum) Gary Voelker, Barrick Museum, Box 454012, University of Nevada Las Vegas, Las Vegas, NV 89154 or (Dept. Biological Sciences) Graduate Coordinator, Dept. of Biological Sciences, University of Nevada Las Vegas, Box 454004, Las Vegas, NV 89154-4004. Telephone: (702) 895-3390.



New Mexico State University, Las Cruces, NM 88003, USA.

Description.--New Mexico State University is located in southern New Mexico. The Department of Fisheries and Wildlife Sciences offers an M .S. degree with emphasis in ornithology. Emphasis is on the biology and management of natural systems, especially of arid lands of the Southwest U. S. and Mexico. There are nine teaching/research faculty in the Department of Fisheries and Wildlife Science, including the U. S. Geological Survey Cooperative Fish and Wildlife Research Unit and Wildlife Extension. There are approximately 170 undergraduate majors and 30 M. S. candidates in the department. The department has a Masters program, and students can train at the Ph.D. level through a variety of other graduate programs on campus. Research projects include big game and predator management, avian ecology, endangered species and conservation biology, fisheries modeling, aquatic toxicology and fish physiology, wetlands management, and habitat characterization using GIS.

Facilities.--New Mexico State University is the site of the New Mexico Fish and Wildlife Cooperative Unit. Students have access to two university "ranches" and to extensive areas of public land in a wide variety of habitats.

Faculty--Members currently involved in ornithological studies are:

  • Craig Benkman,; evolutionary ecology of birds
  • Marv Bernstein,; avian physiology
  • Donald F. Caccamise, Professor and Department head: avian ecology and behavior.
  • F. Graham Cooch, adjunct faculty: waterfowl biology and management.
  • Peter Houde,; avian systematics
  • Bruce C. Thompson, adjunct faculty and New Mexico Cooperative Fish and Wildlife Unit: biodiversity conservation, sustainable resource use, avian ecology.
  • Raul Valdez, Professor: ecological relationships of the Aplomado Falcon, ungulate ecology, behavior, and taxonomy.
For information.--Write to Department of Fishery and Wildlife Sciences, P. O. Box 30003, MSC 4901, New Mexico State University, Las Cruces, NM 88003-8003, or visit

For information on the Graduate School, write: Graduate School, Dept. 3G, New Mexico State University, Las Cruces, NM 88003-8001

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